The Department of Ecology and Evolution

The Department of Ecology and Evolution provides training for research and teaching in the ecology, evolution and behavior of whole organisms, at the levels of the organism, the population, and the ecosystem. The research interests of our faculty include molecular evolution, population genetics, quantitative genetics, animal behavior, plant and animal ecology, evolutionary theory, systematics, paleontology, and related subjects. Individual levels of study range from molecules to communities. A common theme is the conduct of studies in a rigorous ecological and conceptual context, and the faculty share an interest in the architecture of populations, species and communities.

The department stresses scientific breadth and the interrelations between various specialized fields. Students are encouraged to approach basic biological problems with the most appropriate techniques: biophysical, biochemical, mathematical, physiological, or organismal. Departmental laboratories are equipped for a wide variety of contemporary research methods. Courses in other departments may be taken for credit in ecology and evolution—for example, in the Departments of Organismal Biology and Anatomy, Biochemistry and Molecular Biology, Molecular Genetics and Cell Biology, Statistics, Geophysical Sciences, Anthropology, and Chemistry. Many students in the Department of Ecology and Evolution participate in interdepartmental programs in genetics, cell biology, developmental biology, population biology, theoretical biology, and evolutionary biology, and in these programs dissertation research may be cosponsored by faculty from different departments. Collaboration is also maintained with the Field Museum and the Shedd Aquarium for students interested in research in systematics, taxonomy, and evolutionary biology, and with the Brookfield Zoo for basic research in conservation and behavior involving zoo animals. Possibilities also exist for field studies in Central America, Africa, and other regions of the earth.

Program of Study

Most students in the Department of Ecology and Evolution complete their Ph.D. program in about five years, though students entering with master’s degrees may finish in slightly less time. A student advisory committee advises all incoming and second year students on academic and research concerns. The first and second years consist largely of course work and individual reading courses, aiming toward successful completion of an oral general knowledge examination by the spring quarter of the first year, supervised by the student advisory committee and the department chair. The student and faculty advisor then choose a five-member faculty doctoral committee, scheduling a defense of the dissertation research proposal by the end of the second year of study.
Work in subsequent years shifts to dissertation-centered research and, finally, preparation and defense of the Ph.D. dissertation. All students are required to be a teaching assistant in two approved courses during their tenure in the doctoral program. While there is no master’s program in the department, students may elect to receive the S.M. degree upon successful completion of their dissertation proposal defense.

Entrance requirements

Entering students are expected to have received a broad undergraduate training in biology, and a good background in related quantitative subjects, such as chemistry, statistics and calculus. Students who are admitted without having fully satisfied these requirements will be required to remedy their deficiencies by taking appropriate courses during their first two years in the graduate program.

General Knowledge Examination

Each first year student will be expected to pass an oral general knowledge examination during the first year of study, generally no later than the 10th week of the spring quarter. This examination session shall be attended by all three members of an examination committee appointed, in writing, by the department chair; the committee will be composed of two faculty members in the student’s general area of research, and one member of the student advisory committee. The goal of the examination will be to assess each student’s general knowledge of key concepts, processes and issues in ecology and evolutionary biology, as covered in the courses recommended to the student by the student advisory committee during the student’s first year in the program.

Dissertation Proposal Defense

This examination consists of the submission of a written Ph.D. research proposal and an oral presentation of the proposal in a public or closed/private seminar format, followed by a closed discussion and examination on the proposal presentation with the faculty committee chosen by the student and the chair of the department. Students are expected to schedule the dissertation proposal defense before the end of their second year.

Doctor of Philosophy

Upon successful completion of the dissertation proposal defense and admission into candidacy for the Ph.D., students work closely with the faculty advisor and dissertation committee on the dissertation project. During the period of two to three years in which students do primary original research, they also participate in seminars, discussion groups, and professional meetings and conferences, leading to the completion of the written Ph.D. dissertation. The Ph.D. in Ecology and Evolution is awarded based upon (1) submission of a written dissertation based on original research, which must be approved by the faculty adviser and dissertation committee; (2) presentation of a public seminar based on the dissertation research; (3) following the public seminar, successful performance during an oral examination by the dissertation committee and other relevant faculty; and, (4) acceptance of the approved written dissertation by the University Office of Academic Publications in compliance with that office’s regulations.

Application

Applications for admission to the department may be made at any time, though students applying for financial support should submit formal applications no later than January 5. We strongly advise students considering application to the department to begin preparation of their application early in the autumn quarter, so that all materials will arrive by the January 5 deadline. The department requires GRE General Test scores from all applicants, and strongly recommends submission of GRE subject test scores in biology. Foreign applicants whose first language is not English also must submit TOEFL test scores with their application materials.
Interested students should write to the Administrative Director, Graduate Programs, Darwinian Sciences, 1101 East 57th Street, Chicago, IL 60637, for a brochure describing the graduate program and faculty research interests, and to obtain required University application materials. Further information also may be obtained from the department’s home page on the World Wide Web, at http://pondside.uchicago.edu/darwin, or by sending an e-mail to Darwin@pondside.uchicago.edu.

This text was last revised on 9/05/2003.